細看針葉樹(7)–看台灣二葉松的原生葉與次生葉

在台灣木本植物誌裡的松亞科松屬特徵裡提到:

松屬的葉子有兩種型態:原生葉與次生葉

   

 原生葉螺旋狀排列且常退化為小膜質苞

  次生葉針形,橫斷面半圓或三角形著生於短椏上

 

原生葉特寫—深褐色片狀螺旋狀排列於幼芽上

退化為小膜質苞–功能與芽鱗(bud scale)相同

 

 

原生葉腋下會長出短枝及針狀的次生葉,圖中可見短枝的幼芽

 

去除原生葉之後可見數個短椏幼芽披覆深褐色鱗片

 

上圖中可見針狀葉基部開裂的鱗片

 

正常情況下短椏基部的鱗片包裹針狀葉的情形如上

 

這些鱗片好像會永存!

 

這即是短椏及短椏上的針狀葉(也就是次生葉)

 

圖中A 為長椏, B 為短椏

 

短椏圖形參考文獻

Morphology, anatomy and vasculature of leaves in Pinus (Pinaceae) and its evolutionary meaning

PDF

http://www.sciencedirect.com/science/article/pii/S0367253011001563

 

原生葉(primary leaf) 與次生葉(secondary leaf)可參考google 圖書內的內容

Illustrated guide to trees and shrubs: a handbook of the woody … – Google 圖書結果

books.google.com.tw/books?isbn=0486272583Arthur Harmount Graves – 1956 – Nature – 288 頁
The primary leaves are usually small and scale-like and function as bud scales. But in the Pine, in the young seedling stage, the primary leaves are the only leaf

 

 

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